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Phiên bản lúc 07:04, ngày 14 tháng 10 năm 2011

Map of early human migrations according to mitochondrial population genetics (numbers are millennia before present) Bản mẫu:Underdiscussion-inline.

Các cuộc di cư sớm thời tiền sử bắt đầu khi loài Homo erectus di cư lần đầu tiên ra khỏi châu Phi qua hành lang Levantine và Sừng châu Phi tới lục địa Á-Âu khoảng 1,8 triệu năm về trước, cuộc di cư có thể bắt đầu do sự phát triển của ngôn ngữ (hay ngôn ngữ sơ khai theo giả thuyết của Fischer. [1] Việc bành trướng của H. erectus khỏi châu Phi được tiếp nối bởi Homo antecessor vào châu Âu khoảng 800.000 năm trước, tiếp theo là Homo heidelbergensis khoảng 600.000 năm trước, tại đây có thể họ đã tiến hóa thành người Neanderthal.[2]

Người hiện đại, Homo sapiens, tiến hóa tại châu Phi cho tới 200.000 năm trước và tiến vào Cận Đông khoảng 125.000 năm trước.[3] Từ Cận Đông, những người này mở rộng về phía đông tới Nam Á vào khoảng 50.000 năm trước, và tiếp tục tới châu Úc khoảng 40.000 năm trước, [4] khi lần đầu tiên H. sapiens đặt chân đến các vùng đất H. erectus chưa bao giờ đến. H. sapiens tới châu Âu khoảng 40.000 năm trước, và cuối cùng đã thay thế người Neanderthal. Họ tới Đông Á vào khoảng 30.000 năm trước.

The date of migration to North America is disputed; it may have taken place around 30 millennia ago, or considerably later, around 14 millennia ago. Colonisation of the Pacific islands of Polynesia began around 1300 BC, and was completed by 900 AD. The ancestors of Polynesians left Taiwan around 5200 years ago.

Người tiền sử (trước Homo sapiens)

Xem thêm thông tin: Yuanmou Man và Lantian Man
Tập tin:Homo erectus.JPG
A reconstruction of Homo erectus. Anthropologists believe that H. erectus was the first hominid to control fire.

Early members of the Homo genus, i.e. Homo ergaster, Homo erectus and Homo heidelbergensis, migrated from Africa during the Early Pleistocene, possibly as a result of the operation of the Saharan pump, around 1.9 million years ago, and dispersed throughout most of the Old World, reaching as far as Southeast Asia. The date of original dispersal beyond Africa virtually coincides with the appearance of Homo ergaster in the fossil record, and the associated first emergence of full bipedalism, and about half a million years after the appearance of the Homo genus itself and the first stone tools of the Oldowan industry. Key sites for this early migration out of Africa are Riwat in Pakistan (1.9 Mya), Ubeidiya in the Levant (1.5 Mya) and Dmanisi in the Caucasus (1.7 Mya).

China was populated more than a million years ago,[5] as early as 1.66 Mya based on stone artifacts found in the Nihewan Basin.[6] Stone tools found at Xiaochangliang site were dated to 1.36 million years ago.[7] The archaeological site of Xihoudu (西侯渡) in Shanxi Province is the earliest recorded use of fire by Homo erectus, which is dated 1.27 million years ago.[5]

Southeast Asia (Java) was reached about 1.7 million years ago (Meganthropus). West Europe was first populated around 1.2 million years ago (Atapuerca).[8]

Bruce Bower[cần dẫn nguồn] has suggested that Homo erectus may have built rafts and sailed oceans, a theory that has raised some controversy.[9]

Homo sapiens sapiens migrations

Homo sapiens sapiens is supposed to have appeared in East Africa between 200,000 and 100,000 years ago (Homo sapiens idaltu, found at site Middle Awash in Ethiopia, lived about 160,000 years ago,[10] is considered the oldest known anatomically modern human).

From there they spread around the world. An exodus from Africa over the Arabian Peninsula around 60,000 years ago brought modern humans to Eurasia, with one group rapidly settling coastal areas around the Indian Ocean and one group migrating north to steppes of Central Asia.[11]

The migration path is a matter of debate and study. Genetics have shed some light on this matter.

Within Africa

The matrilinear most recent common ancestor shared by all living human beings, dubbed Mitochondrial Eve, probably lived roughly 120-150 millennia ago,[12] the time of Homo sapiens idaltu, probably in East Africa.[cần dẫn nguồn]

The broad study of African genetic diversity headed by Dr. Sarah Tishkoff found the San people to express the greatest genetic diversity among the 113 distinct populations sampled, making them one of 14 "ancestral population clusters." The research also located the origin of modern human migration in south-western Africa, near the coastal border of Namibia and Angola.[13]

Around 100,000-80,000 years ago, three main lines of Homo sapiens diverged. Bearers of mitochondrial haplogroup L0 (mtDNA) / A (Y-DNA) colonized Southern Africa (the ancestors of the Khoisan (Capoid) peoples), bearers of haplogroup L1 (mtDNA) / B (Y-DNA) settled Central and West Africa (the ancestors of western pygmies), and bearers of haplogroups L2, L3, and others mtDNA remained in East Africa (the ancestors of Niger–Congo- and Nilo-Saharan-speaking peoples). (see L-mtDNA)

Exodus from Africa

Bài chi tiết: Recent African origin of modern humans
Red sea crossing

According to the Recent African Origin hypothesis a small group of the L3 bearers living in East Africa migrated north east, possibly searching for food or escaping adverse conditions, crossing the Red Sea about 70 millennia ago, and in the process going on to populate the rest of the world. According to some authors, based in the fact that only descents of L3 are found outside Africa, only a few people left Africa in a single migration to a settlement in the Arabian peninsula.[14] From that settlement, some others point to the possibility of several waves of expansion close in time. For example, Wells says that the early travelers followed the southern coastline of Asia, crossed about 250 kilometers [155 miles] of sea (probably by simple boats or rafts[15]), and colonized Australia by around 50,000 years ago. The Aborigines of Australia, Wells says, are the descendants of the first wave of migration out of Africa.[16]

Around 50,000 years ago the world was entering the last ice age and water was trapped in the polar ice caps, so sea levels were much lower. Today at the Gate of Grief the Red Sea is about 12 miles (20 kilometres) wide but 50,000 years ago it was much narrower and sea levels were 70 metres lower. Though the straits were never completely closed, there may have been islands in between which could be reached by simple rafts. Shell middens 125,000 years old indicate that the diet of early humans in Eritrea included sea food obtained by beachcombing. This has been seen as evidence that humans may have crossed the Red Sea in search of food sources on new beaches.

South Asia and Australia

Xem thêm: Prehistory of Australia và South Asian Stone Age

Some genetic evidence points to migrations out of Africa along two routes. However, other studies suggest that a single migration occurred, followed by rapid northern migration of a subset of the group. Once in West Asia, the people who remained south (or took the southern route) spread generation by generation around the coast of Arabia and Persia until they reached India. One of the groups that went north (east Asians were the second group) ventured inland[17] and radiated to Europe, eventually displacing the Neanderthals. They also radiated to India from Central Asia. The former group headed along the southeast coast of Asia, reaching Australia between 55,000 and 30,000 years ago,[4] with most estimates placing it about 46,000 to 41,000 years ago.

The map shows the probable extent of land and water at the time of the last glacial maximum, 20,000 yrs BP and when the sea level was probably more than 110m lower than today

During that time, sea level was much lower and most of Maritime Southeast Asia was one land mass known as the lost continent of Sunda. The settlers probably continued on the coastal route southeast until they reached the series of straits between Sunda and Sahul, the continental land mass that was made up of present-day Australia and New Guinea. The widest gaps are on the Weber Line and are at least 90 km wide,[18] indicating that settlers had knowledge of seafaring skills. Archaic humans such as Homo erectus never reached Australia, although they crossed the Lombok gap reaching as far as Flores.

If these dates are correct, Australia was populated up to 10,000 years before Europe. This is possible because humans avoided the colder regions of the North favoring the warmer tropical regions to which they were adapted given their African homeland. Another piece of evidence favoring human occupation in Australia is that beginning about 46,000 years ago, all megafauna weighing more than 100 kg became extinct. Tim Flannery and others argue new settlers were likely to be responsible for this extinction.[19] Many of the animals may have been accustomed to living without predators and become docile and vulnerable to attack (as occurred later in the Americas).

While some settlers crossed into Australia, others may have continued eastwards along the coast of Sunda eventually turning northeast to China and finally reaching Japan, leaving a trail of coastal settlements. This coastal migration leaves its trail in the mitochondrial haplogroups descended from haplogroup M, and in Y-chromosome haplogroup C. Thereafter, it may have become necessary to venture inland possibly bringing modern humans into contact with archaic humans such as H. erectus. Recent genetic studies suggest that Australia and New Guinea were populated by one single migration from Asia as opposed to several waves. The land bridge connecting New Guinea and Australia became submerged approximately 8,000 years ago, thus isolating the populations of the two land masses.[20][21]

Europe

Bài chi tiết: Paleolithic Europe
Bài chi tiết: Neolithic Europe

Europe is thought to have been colonized by northwest bound migrants from Central Asia and the Middle East. When the first anatomically modern humans entered Europe, Neanderthals were already settled there. Debate exists whether modern human populations interbred with Neanderthal populations, most of the evidence suggesting that it happened to a small degree rather than complete absorption. Populations of modern man and Neanderthal overlapped in various regions such as in Iberian peninsula and in the Middle East and that interbreeding may have contributed Neanderthal genes to palaeolithic and ultimately modern Eurasians and Oceanians.

An important difference between Europe and other parts of the inhabited world was the northern latitude. Archaeological evidence suggests humans, whether Neanderthal or Cro-Magnon, reached sites in Arctic Russia by 40,000 years ago.[22]

Around 20,000 BC, 10,000 years after the Neanderthal extinction, the Last Glacial Maximum took place forcing northern hemisphere inhabitants to migrate to several shelters until the end of this period came. The resulting populations, whether interbred with Neanderthals or not, are then presumed to have resided in those hypothetical refuges during the LGM to ultimately reoccupy Europe where archaic historical populations are considered their descendents. An alternate view is that modern European populations have descended from Neolithic populations in the Middle East that have been well documented in this area. The debate surrounding the origin of Europeans has been worded in terms of cultural diffusion versus demic diffusion. Archeological evidence and genetic evidence strongly support demic diffusion, that a population spread from the Middle East over the last 12,000 years. A scientific genetic concept called the Time to Most Recent Common Ancestor or TMRCA has been used to refute the demic diffusion in favour of cultural diffusion.[23]

Migration of the Cro-Magnons into Europe

Bài chi tiết: Cro-Magnon

Cro-Magnon are considered the first anatomically modern humans in Europe. They entered Eurasia by the Arabian Peninsula around 60,000 years ago, with one group rapidly settling coastal areas around the Indian Ocean and one group migrating north to steppes of Central Asia.[11]

A mitochondrial DNA sequence of two Cro-Magnons from the Paglicci Cave in Italy, dated to 23,000 and 24,000 years old (Paglicci 52 and 12), identified the mtDNA as Haplogroup N, typical of the latter group.[24] The inland group is the founder of both North- and East Asians (the "Mongol" people), Caucasoids and large sections of the Middle East and North African population. Migration from the Black Sea area into Europe started some around 45,000 years ago, probably along the Danubian corridor. By 20,000 years ago, the whole of Europe was settled. The first complete human remains that have been preserved is the mummy Ötzi, from 3000 BC, belonging to the K1 MitDNA.

Migration of modern humans into Europe, based on simulation by Currat & Excoffier (2004)[25]
 (YBP=Years Before Present)
Up to 37 500 YBP
Up to 35 000 YBP
Up to 32 500 YBP
Up to 30 000 YBP

Competition with Neanderthals

The expansion is thought to have begun 45,000 years ago and may have taken up to 15,000 years for Europe to be colonized.[17][26]

During this time the Neanderthals were slowly being displaced. Because it took so long for Europe to be occupied, it appears that humans and Neanderthals may have been constantly competing for territory. The Neanderthals were larger and had a more robust or heavy built frame which may suggest that they were physically stronger than modern Homo sapiens. Having lived in Europe for 200,000 years they would have been better adapted to the cold weather. The anatomically modern humans known as the Cro-Magnons, with superior technology and language would eventually completely displace the Neanderthals, whose last refuge was in the Iberian peninsula. After about 30,000 years ago the fossil record of the Neanderthals ends, indicating that they had become extinct. The last known population lived around a cave system on the remote south-facing coast of Gibraltar from 30,000 to 24,000 years ago.

Proponents of the multiregional hypothesis have long believed that Europeans were descended from Neanderthals and not from this homo-sapiens migration. Others believed the Neanderthals had interbred with modern humans. In 1997 researchers managed to extract mitochondrial DNA from a 40,000 year old specimen of a Neanderthal. On comparison with human DNA, its sequences differed significantly, indicating that based on the mitochondrial DNA, modern Europeans are not descended from the Neanderthals and that no interbreeding took place.[27] Some scientists continue to search autosomal DNA for traces of Neanderthal admixture.[28] A few alleles of some autosomal genes such as the H2 allele of the MAPT gene have been suggested, since they were only found among Europeans. In the absence of autosomal DNA from a Neanderthal, the scientists conclude that this hypothesis is speculative.[29]

Some archaeologists suspect that Neanderthals and Homo sapiens were not interfertile. This is because Neanderthals and Europeans shared the same habitat for up to 20,000 years, yet no undisputed skeletal fossils have been found that show intermediate properties between the two species.[30]

Current (as of 2010) genetic evidence suggests interbreeding took place with Homo sapiens sapiens (anatomically modern humans) between roughly 80,000 to 50,000 years ago in the Middle East, resulting in non-ethnic sub-Saharan Africans having no Neanderthal DNA and Caucasians and Asians having between 1% and 4% Neanderthal DNA.[31]

Central and Northern Asia

Mitochondrial haplogroups A, B and G originated about 50,000 years ago, and bearers subsequently colonized Siberia, Korea and Japan, by about 35,000 years ago. Parts of these populations migrated to North America.

Americas

Bài chi tiết: Settlement of the Americas và Genetic history of indigenous peoples of the Americas
A map of early human migrations[32]

The specifics of Paleo-Indians migration to and throughout the American Continent, including the exact dates and routes traveled, are subject to ongoing research and discussion.[33] The traditional theory has been that these early migrants moved into the Beringia land bridge between eastern Siberia and present-day Alaska around 40,000 — 17,000 years ago,[34] when sea levels were significantly lowered due to the Quaternary glaciation.[33][35] These people are believed to have followed herds of now-extinct pleistocene megafauna along ice-free corridors that stretched between the Laurentide and Cordilleran ice sheets.[36] Another route proposed is that, either on foot or using primitive boats, they migrated down the Pacific Northwest coast to South America as far as Chile.[37] Evidence of the latter would since have been covered by a sea level rise of a hundred meters following the last ice age.[38]

Archaeologists contend that Paleo-Indians migration out of Beringia (eastern Alaska), ranges from 40,000 to around 16,500 years ago.[39][40][41] This time range is a hot source of debate and will be for years to come. The few agreements achieved to date are the origin from Central Asia, with widespread habitation of America during the end of the last glacial period, or more specifically what is known as the late glacial maximum, around 16,000 — 13,000 years before present.[34][42]

See also

References

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  2. ^ Finlayson, Clive (2005). “Biogeography and evolution of the genus Homo”. Trends in Ecology & Evolution. Elsevier. 20 (8): 457–463. doi:10.1016/j.tree.2005.05.019. PMID 16701417.
  3. ^ “Hints Of Earlier Human Exit From Africa”. Science News. doi:10.1126/science.1199113. Kiểm tra giá trị |doi= (trợ giúp). Truy cập ngày 1 tháng 5 năm 2011.
  4. ^ a b Bowler, James M. et al 2003, New ages for human occupation and climatic change at Lake Mungo, Australia
  5. ^ a b Rixiang Zhu, Zhisheng An, Richard Pott, Kenneth A. Hoffman (2003). “Magnetostratigraphic dating of early humans in China” (PDF). Earth Science Reviews. 61 (3–4): 191–361. Bibcode:2003ESRv...61..191A. doi:10.1016/S0012-8252(02)00110-1. Đã bỏ qua tham số không rõ |month= (trợ giúp)Quản lý CS1: nhiều tên: danh sách tác giả (liên kết)
  6. ^ R. Zhu et al. (2004), New evidence on the earliest human presence at high northern latitudes in northeast Asia.
  7. ^ “Earliest Presence of Humans in Northeast Asia”. Human Origins Program. Smithsonian Institution. Truy cập ngày 4 tháng 8 năm 2007.
  8. ^ Hopkin M (26 tháng 3 năm 2008). “'Fossil find is oldest European yet'”. Nature News. doi:10.1038/news.2008.691.
  9. ^ Bednarik RG (2003). “Seafaring in the Pleistocene”. Cambridge Archaeological Journal. 13 (1): 41–66. doi:10.1017/S0959774303000039.
    ScienceNews summary
  10. ^ White, Tim D., Asfaw, B., DeGusta, D., Gilbert, H., Richards, G.D., Suwa, G. and Howell, F.C. (2003). “Pleistocene Homo sapiens from Middle Awash, Ethiopia”. Nature. 423 (6491): 742–747. doi:10.1038/nature01669. PMID 12802332.Quản lý CS1: nhiều tên: danh sách tác giả (liên kết)
  11. ^ a b “Atlas of human journey: 45 - 40,000”. The genographic project. National Geographic Society. 1996–2010. Truy cập ngày 31 tháng 3 năm 2010. |chapter= bị bỏ qua (trợ giúp)Quản lý CS1: định dạng ngày tháng (liên kết)
  12. ^ Misconceptions Around Mitochondrial Eve, Alec MacAndrew.
  13. ^ BBC World News "Africa's genetic secrets unlocked", 1 May 2009; the results were published in the online edition of the journal Science.
  14. ^ “Both Australian Aborigines and Europeans Rooted in Africa”. News.softpedia.com. Truy cập ngày 11 tháng 1 năm 2011.
  15. ^ Evolution of modern humans, truy cập ngày 25 tháng 12 năm 2010
  16. ^ “Human line 'nearly split in two'”. BBC News. 24 tháng 4 năm 2008. Truy cập ngày 31 tháng 12 năm 2009.
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  18. ^ “Pleistocene Sea Level Maps”. Fieldmuseum.org. Truy cập ngày 23 tháng 9 năm 2010.
  19. ^ Flannery, Tim (2002), "The Future Eaters: An Ecological History of the Australasian Lands and People" (Grove Press)
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  22. ^ Pavlov, Pavel (2001). “Human presence in the European Arctic nearly 40,000 years ago”. Nature. 413 (6851): 64–67. doi:10.1038/35092552. PMID 11544525. Đã bỏ qua tham số không rõ |month= (trợ giúp); Đã bỏ qua tham số không rõ |coauthors= (gợi ý |author=) (trợ giúp)
  23. ^ “A Comparison of Y-Chromosome Variation in Sardinia and Anatolia Is More Consistent with Cultural Rather than Demic Diffusion of Agriculture”. Plosone.org. Truy cập ngày 23 tháng 9 năm 2010.
  24. ^ Caramelli, D; Lalueza-Fox, C; Vernesi, C; Lari, M; Casoli, A; Mallegni, F; Chiarelli, B; Dupanloup, I; Bertranpetit, J; Barbujani, G; Bertorelle, G (2003). “Evidence for a genetic discontinuity between Neandertals and 24,000-year-old anatomically modern Europeans” (Free full text). Proceedings of the National Academy of Sciences of the United States of America. 1 (11): 6593–7. Bibcode:2003PNAS..100.6593C. doi:10.1073/pnas.1130343100. ISSN 0027-8424. PMC 164492. PMID 12743370. Đã bỏ qua tham số không rõ |month= (trợ giúp)Quản lý CS1: nhiều tên: danh sách tác giả (liên kết)
  25. ^ Currat, M. & Excoffier, L. (2004): Modern Humans Did Not Admix with Neanderthals during Their Range Expansion into Europe. PLoS Biol no 2 (12): e421. doi:10.1371/journal.pbio.0020421
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  38. ^ “68 Responses to "Sea will rise 'to levels of last Ice Age'"”. Center for Climate Systems Research, Columbia University. Truy cập ngày 17 tháng 11 năm 2009.
  39. ^ “Introduction”. Government of Canada. Parks Canada. 2009. Truy cập ngày 9 tháng 1 năm 2010. Canada's oldest known home is a cave in Yukon occupied not 12,000 years ago like the U.S. sites, but at least 20,000 years ago
  40. ^ “Pleistocene Archaeology of the Old Crow Flats”. Vuntut National Park of Canada. 2008. Truy cập ngày 10 tháng 1 năm 2010. However, despite the lack of this conclusive and widespread evidence, there are suggestions of human occupation in the northern Yukon about 24,000 years ago, and hints of the presence of humans in the Old Crow Basin as far back as about 40,000 years ago.
  41. ^ “Jorney of mankind”. Brad Shaw Foundation. Truy cập ngày 17 tháng 11 năm 2009.
  42. ^ “A single and early migration for the peopling of greater America supported by mitochondrial DNA sequence data”. The National Academy of Sciences of the US. National Academy of Sciences. Truy cập ngày 10 tháng 10 năm 2009.

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